The Times assistant editor Andrew Pierce is to join the Daily Telegraph in the same role.
Pierce, 45, joins existing Daily Telegraph assistant editor Iain Martin, who moved from the editorship of Scotland on Sunday in March.
Pierce may now be a contender for the vacant editor's chair. Telegraph Group editor-in-chief John Bryant has been running the Daily Telegraph since the resignation of editor Martin Newland in November 2005.
After beginning his journalism career in local newspapers and then freelancing for the nationals, Pierce first joined The Times in 1988. Following a spell as assistant editor at the Daily Express, he rejoined The Times in 1999, also as assistant editor.
Pierce said: "I am delighted to be joining the leading quality daily newspaper at such an exciting time in its development."
Bryant said: "Andrew is a brilliant journalist, with a proven ability to uncover important and newsworthy stories. He'll be an enormous asset to the paper."